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Some commonly asked questions and interesting facts about spiders.


Are huntsman spiders dangerous? They look so large and hairy...

Despite their often large and hairy appearance, huntsman spiders are not considered to be dangerous spiders. As with most spiders, they do possess venom, and a bite may cause some ill effects. However, they are quite reluctant to bite, and will usually try to run away rather than be aggressive. In houses they perform a useful role as natural pest controllers.

Some people may think of huntsman spiders as 'tarantulas'. However, they are not related to the large hairy ground dwelling spiders that are normally called tarantulas. Both huntsman spiders and tarantulas are often portrayed as being dangerous and scary. This usually is the case in films or stories that deliberately present spiders in a frightening and unrealistic way. If you feel frightened of huntsman spiders because of this, perhaps you might like to learn more about their true habits and biology. In this way you might be able to reduce your fears.


Huntsman
Huntsman spider with an eggsac Image: Stuart Humphreys
© Australian Museum

How do you identify a wolf spider?

One of the diagnostic features of wolf spiders is their eye pattern which comprises three rows at the front of the carapace: four (smaller) eyes in the first row, two above the first and two above the second row. The diagram below (basically) shows this layout, face-on to the spider:

top of the head

0 0

0 0

. . . .

bottom of the head

Wolf spiders also have a variegated pattern on their bodies, often including radiating lines on the carapace and scroll-like patterns on the top of the abdomen. The underside of the spider is grey or black, sometimes with white markings. They can have orange spots on the sides of their jaws.

As wolf spiders actively hunt for food they are likely to be found roving along the ground and they are more active at night. When spotlighted at night wolf spider's eyes will glow green. Scientists use this method during invertebrate surveys.


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Does Australia have a bird-eating spider?

The term 'bird-eating spider' usually refers to large spiders from the family Theraphosidae. These spiders are also referred to as tarantulas. In Australia the theraphosids are represented by the whistling spiders (Selenocosmia sp.). These ground-dwelling spiders are big enough to prey on small frogs and reptiles, but are not known to eat birds. They are also known as barking spiders.

Do we have tarantulas in Australia?

It depends on what you mean by the word "tarantula". Some people use it to describe the large hairy spiders of South and Central America. In Australia, the whistling spiders are also called Australian tarantulas, as they are related to the American spiders. However, the word tarantula is also used to refer to huntsman spiders.

Tarantula is derived from the name of a town in Italy, Taranto. This town is the original home of the wild dance called the tarentella. During the Middle Ages, the tarentella was thought to be the way to cure the bite of a particular spider. The symptoms - known as tarantism - included severe pain, swelling, spasms, nausea and vomiting, palpitations, and fainting, along with exhibitionism, melancholia and delirium. It was hard to determine whether an actual bite had occurred or if people were merely displaying some form of madness or hysteria. Scientists later determined that many cases might indeed have been the result of a bite, although much of the fierce dancing and extreme behaviour may reflect more about the social and sexual repression at the time.

The alleged spider that caused all of these symptoms was called a tarantula, but the species was incorrectly identified. The original spider identified by the people of the time was a wolf spider (Lycosa tarantula). However, it was subsequently shown to cause little serious results when it bit people. Finally, it was shown that the real culprit was a Black Widow relative, Latrodectus tredecimguttatus, known in Southern Europe as the "malmignatte". The symptoms of this spider's bite (and of other Latrodectus species, including the Redback Spider) match the whole-body symptoms experienced during tarantism.

Information from: Hillyard, P. 1994. The Book of the Spider. Hutchinson, London.

Do we have scorpions in Australia?

Yes we do. Scorpions are common in gardens and forests throughout eastern Australia and are found under logs, rocks and in shallow burrows in earth banks. They are nocturnal - which is why we rarely see them - but they can be disturbed during the day, especially during the prolonged wet weather. There are also species that live in the desert and others that inhabit tropical rainforests.

What is the world's most dangerous spider?

It is hard to define which spider in the world is the most dangerous to humans. Several spiders could qualify, depending on what you mean by dangerous. Do you mean the spider with the most toxic venom, measured by its effect on newborn mice or other mammals? Or do you mean the spider that has caused the death of the most people? Those that have the strongest venom may not be encountered by humans very often, or may even have trouble piercing human skin and so are not considered to be 'dangerous'. Data are usually only kept on bites from spiders that are potentially deadly or cause severe reactions and these data are not recorded consistently at a national or international level. Here, we will define dangerous as 'deadly'.

In summary, on current evidence the most dangerous spiders in the world are funnel-web spiders (Atrax and Hadronyche species), Redback Spiders and their relations (Latrodectus species), Banana Spiders (Phoneutria species) and Recluse Spiders (Loxosceles species). In Australia, only male Sydney Funnel Web Spiders and Redback Spiders have caused human deaths, but none have occurred since antivenoms were made available in 1981.

The Australian funnel-web spiders are among the deadliest spiders in the world in the effect their bites have on humans and our primate relations (although the bite has little effect on dogs and cats). There are many species of funnel-web spiders in Australia but only male Sydney Funnel-webs have caused human deaths. There have been only 13 deaths recorded from male Sydney Funnel-webs, but up to 30-40 people are bitten by funnel-web spiders each year. Mouse spiders may have venom that is as toxic as that of some funnel-webs, as some patients have had severe reactions to their bites, although no-one has been recorded as having died from the effects of a mouse spider bite. Antivenoms are available for both funnel-web and Redback Spider bites.

A group of spiders that is dangerous in many countries belongs to the genus Latrodectus in the Family Theridiidae. In Australia we have the Redback Spider (Latrodectus hasselti). In America, a common representative of this genus is the Black Widow (Latrodectus mactans). Antivenoms are available for both funnel-web and Redback Spider bites.

A deadly spider which comes from South America is the Banana Spider, Phoneutria species. In south-eastern Brazil between 1970 and 1980, more than 7,000 people were admitted to hospital with bites from this spider. An antivenom also exists for this species.

The Recluse or Fiddleback Spider is a deadly spider belonging to the genus Loxosceles. Recluse spiders are found in many parts of the world and have been introduced into Australia. The venom of this spider can cause severe skin necrosis (eating away of the flesh) and can be fatal although not many deaths have been recorded.

How many dangerous spider bites occur in Australia each year? Has anyone died from a bite recently?

There have been no deaths in Australia from a confirmed spider bite since 1979. An effective antivenom for Redback Spiders was introduced in 1956, and one for funnel-web spiders in 1980. These are the only two spiders that have caused deaths in Australia in the past.

A spider bite is not a notifiable medical emergency, so there are no Australia-wide statistics, but the following figures give an idea of the incidence of reported bites in recent years.

Approximately 2000 people are bitten each year by Redback Spiders

Funnel-web spider antivenom has been given to at least 100 patients since 1980. Antivenom is given only when signs of serious envenomation are observed. Many spider bites are 'blank', which means that no venom has been injected.

During 2000 the New South Wales Poisons Information Centre received 4,200 calls about spiders. However not all of these would have involved actual bites. Many reported bites are not able to be identified as definitely being from a spider, and it is nearly impossible to work out what species has caused a bite without seeing a specimen of the spider responsible.

Figures are from: Sutherland, S K and Nolch, G (2000) Dangerous Australian Animals. Hyland House, Flemington, Vic. 201 pp. ISBN 86447 076 3

  • Poisons Information Centre
  • The Children's Hospital at Westmead
  • Locked Bag 4001
  • Westmead, NSW 2145
  • Emergency telephone: 131 126 (24 hours, within Australia only)
  • Administrative telephone: +61 2 9845 3111
  • Fax: +61 2 9845 3597

What spiders in Australia may cause ill effects if they bite you?

In Australia, bites from at least two kinds of spiders - wolf spiders and white-tailed spiders - in some cases cause skin necrosis (eating away of the flesh). However, neither spider has caused human deaths. There are also a number of others which are thought to cause the same problem, but research is still being done to find out exactly which species do so.

Bites from many Australian spiders can cause localised reactions, with symptoms such as swelling and local pain at the site of the bite, sweating, nausea and vomiting and headaches. All of these symptoms will vary in severity depending on the age of the victim, their health, and the amount of venom that the spider was able to inject. Have a look at our spider fact sheets to find out more about individual species.

Do white-tailed spiders cause the skin condition known as necrotising arachnidism?

There is an ongoing debate among toxicologists and spider biologists about the effects and dangers of white-tailed spider bites. Most of these bites appear to cause little or no effect beyond transient local pain. However a small number of cases do cause more extensive problems. Whether this is a result of the spiders' venom or to bacteria infecting the wound at or after the time of the bite has not yet been resolved. It is also possible that some people may react badly to white-tailed spider bite, possibly because of immune system susceptibility or a predisposing medical condition.

References

  • Meier, J. & White, J. (1995) Handbook of Clinical Toxicology. CRC Press, Florida USA.
  • Whitehouse, R. (ed.) (1991) Australia's Dangerous Creatures, Readers Digest Pty Ltd, Surry Hills NSW.
  • Sutherland, S. & Sutherland, J. (1999) Venomous Creatures of Australia, Oxford University Press, South Melbourne.
  • Isbister,G. & Greay,M. (2000). "Acute and recurrent skin ulceration after spider bite" Medical Journal of Australia 172, 20 March 2000, pp.303-304

How do I control white-tailed spiders around the house?

Beyond killing or removing all white-tailed spiders that you encounter, you can try a prey reduction strategy. White-tailed spiders like to feed on Black House Spiders (Badumna insignis) in particular, but will take other spiders too. This means that you should clean up obvious spiders around the house (outside and in). This involves removing spiders from around windows, walls and verandas (by web removal and/or direct pyrethrum spray). The condition of the roof cavity and the underfloor area (if raised) should also be investigated. (from Mike Gray, Arachnologist, Australian Museum)

What is the biggest spider in the world?

The biggest spider in the world is the Goliath Spider, Theraphosa leblondi. It lives in coastal rainforests in northern South America. Its body can grow to 9 cm in length (3.5 inches) and its leg span can be up to 28 cm (11 inches). (from: Carwardine, M. 1995. The Guinness Book of Animal Records. Guinness Publishing.)

What is the biggest spider in Australia?

Australia's biggest spiders belong to the same family as the Goliath Spider. They are the whistling spiders. The northern species Selenocosmia crassipes can grow to 6 cm in body length with a leg span of 16 cm.

What is a Daddy-long-legs?

'Daddy-long-legs' is the common name for a particular group of spiders, but it is also used for a different group of arachnids - the harvestmen or opilionids. As a result, there is a lot of confusion about what people mean when they say 'daddy-long-legs'.

Daddy-long-legs: spiders

The animal which most biologists call Daddy-long-legs, is a spider, Pholcus phalangioides, which belongs to the spider family Pholcidae, order Araneida, class Arachnida. It has two parts to the body, separated by a narrow waist. It has eight eyes and eight very long thin legs. Pholcids often live in webs in the corners of houses, sometimes in bathrooms. Daddy-long-legs spiders (or pholcids) kill their prey using venom injected through fangs. Digestion is external, with fluids being squirted onto the prey item and the resulting juices sucked up by the spider.

Daddy-long-legs: harvestmen

The other eight-legged invertebrates that are sometimes called Daddy-long-legs, are members of the order Opiliones or Opilionida in the class Arachnida. Another common name for these arachnids is 'harvestmen'. Unlike spiders, their bodies do not have a 'waist', they do not produce silk and they normally have only one pair of eyes. They do not have venom glands or fangs, although they may produce noxious defence secretions. Most harvestmen eat smaller invertebrates but some eat fungi or plant material and others feed on carcasses of dead mammals and birds. Digestion is internal and some solid food is taken in, which is uncharacteristic for arachnids. You usually do not find harvestmen inside houses.

Are Daddy-long-legs the most venomous spiders in the world?

There is no evidence in the scientific literature to suggest that Daddy-long-legs spiders are dangerously venomous. Daddy-long-legs have venom glands and fangs but their fangs are very small. The jaw bases are fused together, giving the fangs a narrow gape that would make attempts to bite through human skin ineffective.

However, Daddy-long-legs Spiders can kill and eat other spiders, including Redback Spiders whose venom can be fatal to humans. Perhaps this is the origin of the rumour that Daddy-long-legs are the most venomous spiders in the world. The argument is sometimes put that if they can kill a deadly spider they must be even more deadly themselves. However this is not correct. Behavioural and structural characteristics, such as silk wrapping of prey using their long legs, are very important in the Daddy-long-legs' ability to immobilise and kill Redbacks. Also, the effect of the Daddy-long-legs' venom on spider or insect prey has little bearing on its effect in humans.

What are banana spiders and where are they found?

Banana spider is the common name given to large (3 cm body length) active hunting spiders of the genus Phoneutria (Family: Ctenidae). These spiders live in Central and South American rainforests. They are often found in rubbish around human dwellings, as well as hiding in foliage such as banana leaves where they sometimes bite workers harvesting bananas. They have a reputation for being quite aggressive.

Other names for this spider include: Kammspinne, Bananenspinne, Wandering spider, and Aranha armadeira.

The venom of this spider is neurotoxic - acting on the nervous system - and causes little skin damage. Symptoms of a bite include immediate pain, cold sweat, salivation, priapism, cardiac perturbations and occasional death. Research suggests it is similar in action to a-latrotoxin, which is produced by spiders of the Family Latrodectidae, such as the Redback and Black Widow Spiders.

Another spider that seems to have been given the common name "banana spider" is actually a completely unrelated species of orb weaving spider from Florida. This is a good example of why it is more useful to use scientific names when you are trying to find information on different animals or plants.

How do I find out about spiders in New Zealand?

The following New Zealand arachnologist (spider biologist) has offered to respond to inquiries from people interested in New Zealand spiders:

Dr Phil Sirvid
Entomology Section
Museum of New Zealand Te Papa Tongarewa
PO Box 467
Wellington, New Zealand
ph: +644 381 7362
fax: +644 381 7310

There is a book on New Zealand spiders: Forster, Ray and Lyn. 1999. Spiders of New Zealand and Their Worldwide Kin. University of Otago Press, ISBN 1 877133 79 5

What about white-tailed spiders in New Zealand?

Dr Phil Sirvid has this to say about white-tailed spiders in New Zealand:

"We have two species of white-tails in New Zealand - Lampona cylindrata and Lampona murina. They are both very similar in appearance, and can really only be separated from one another by viewing them under a microscope and examining certain features that aren't apparent to the naked eye.

Both have been introduced from Australia.

L. murina has been in the North Island of New Zealand for [over] 100 years, and has also been introduced to the Kermadecs, Lord Howe Island and Norfolk Island. I wouldn't be surprised if it's in the Chatham Islands as well. In Australia, this species is recorded along the East Coast from northern Queensland down through New South Wales and Victoria.

L. cylindrata had only been found occasionally in the South Island until the 1980s. About this time it seemed to spread rapidly throughout the South Island's main urban centres, and is known to occur as far south as Dunedin. This species is found along the southern part of Australia from Western Australia, through South Australia, Victoria and Tasmania, as well as in New South Wales and Queensland."

How can I find out about spiders in North, South or Central America?

We do not have a scientist at the Australian Museum who is an expert on the spiders of the Americas. However you could look at some US spider web sites to see if they can help you. Or you could contact an American spider expert.


Spider biology facts

What is the function and origin of silk glands and spinnerets in spiders?

The development of spinnerets and silk represents a major evolutionary shift that has defined the biological and ecological uniqueness of spiders within the arachnids. Silk glands produce the silk that the spider uses for a variety of purposes. The spinnerets are the special organs that the spider uses to extract and manipulate the silk as is it is produced from the silk glands.

Spiders evolved from ancestors that had limbs on the abdomen, as did arthropods like crustaceans such as crayfish. In fact, one of their few living marine relatives, Limulus, the so-called 'king crabs', has retained abdominal limbs, which have been lost or greatly modified in terrestrial spiders and other arachnids. The spiders' spinnerets are almost certainly derived from these ancestral abdominal limbs. In the basal (lowest) segments of spiders' limbs are small excretory glands - the coxal glands - that secrete and excrete waste body fluids. It seems that the silk glands may represent highly modified excretory glands that now manufacture silk instead of waste products, just as the spinnerets represent highly modified limbs.

It is possible that an intermediate stage in this process could have been the production of a secretion that included pheromone (scent) chemicals put out by the spider as a primitive 'signal line' by which a spider could find its way back to its retreat burrow. This role was then taken over by the production of silk. The silk then became useful not only as a safety line, but also for prey capture, manufacturing egg sacs and a host of other activities.

[Modified from text by Dr Mike Gray - Principal Research Scientist (Spiders)]

Reference: Foelix, R.F.1996. Biology of Spiders. Oxford Thieme.

Why don't spiders get stuck to their webs like the insects that they catch?

If you look at an orb-weaving spider in its web, you'll notice that the body is held slightly clear of the web, especially when the spider is moving about. The spider has only minimal (but vital) body contact with its web via the claws and bristles at the tip of each leg. Compared to its prey, which crashes or blunders into the web, the spider has only a tiny portion of its surface area in contact with a very small amount of silk at any time. This is obviously an important factor when moving on a sticky web - the less contact the better.

Another important factor is that not all silk lines in a sticky web are sticky. For example, the central part of an orb web (where the spider sits) is made of dry silk, as are the spokes supporting the sticky spiral line, which the spider can use when moving around its web. It's only when the spider makes a quick, direct charge across the sticky spiral to capture prey that it may cause some disruption to the web - but it never gets stuck.

Spiders also spend a lot of time grooming their legs. The spider draws the ends of its legs through its jaws to clean them of debris, which may include silk fragments. This is a very important maintenance activity that contributes to efficient function of the claws and bristles. As well as cleaning them, some secretions from the mouthparts may help make the leg tips less susceptible to sticking.

Why don't orb weavers and other spiders fall off their webs?

Most web-building spiders have three claws on their tarsi (feet) - two combed main claws and a smooth central hook. The web silk is only grasped by the hook, and is pushed against serrated bristles, which snag the silk and hold it. When the hook is released by a special muscle, the elastic silk simply springs away from the hook.

Why can some spiders climb slippery surfaces such as glass or run across ceilings?

Many hunting spiders possess dense hair tufts called scopulae under the claws of their tarsi (feet). These scopulae allow many spiders to walk on smooth vertical surfaces, across ceilings and even window panes. Each individual scopula hair splits into thousands of tiny extensions known as end feet. These end feet increase the number of contact points of the tarsi with the surface, creating great adhesion. This is similar to the adhesion forces at work in vertebrates such as skinks and geckos, which can also walk on ceilings with ease. The scopulae can be erected or laid flat by hydraulic pressure through changes in the pressure of the hemolymph (blood supply).

Do spiders sleep?

It really depends on how you define 'sleep'. All animals have some sort of 'circadian' rhythm - a daily activity/inactivity pattern. Some are active during the day - diurnal - others are active at night time - nocturnal/crepuscular. The periods of inactivity are characterised by withdrawal (to a shelter perhaps) and a drop in metabolic rate.

This applies to spiders as well, although no studies have been done to measure the period of time spent in such a state or at what times different species do it. It seems that spiders with good eyesight that rely on vision to capture prey may tend to be more active in daylight hours, whereas others that rely on snares/webs could be active at other times, but this is not necessarily the case for all species.

In cold climates, spiders 'overwinter', which means that they have a kind of hibernation period. Overwintering involves a drop in metabolic rate, where the spiders bring their legs into their body and remain huddled in a shelter during the coldest months of the year.

This ability to shut down for a long period of time indicates that they might be able to do it for shorter periods in their everyday cycle, which could be seen as a form of sleep or rest.

Information from: Foelix, R.F. 1996. Biology of Spiders. Oxford Thieme and the Arachnology section, Australian Museum.


Spider
Jumping Spider, , Portia fimbriata, captured with an LEO Electron Microscopy Ltd. Image: Sue Lindsay
© Australian Museum

Further reading

  • Austin, A.D, and Anderson D.T. (1978). Reproduction and development of the spider Nephila edulis (Koch) (Araneidae: Araneae). Australian Journal of Zoology 26: 501-518.
  • Austin, A.D. and Blest, D.A. (1979). The biology of two Australian species of dinopid spider. Journal of Zoology, London 189: 145-156.
  • Beattie, A. and Ehrlich, P.R. (2000). Wild Solutions: How biodiversity is money in the bank. Melbourne University Press.
  • Bradley, R.A. (1993). Seasonal activity patterns in Sydney funnel-web spiders (Atrax spp. (Araneae: Hexathelidae). Bulletin of the British Arachnological Society 9 (6): 189-192.
  • Clyne, D. (1967). Note on the construction of the net and sperm web of the cribellate spider Dinopis subrufus (Koch) (Araneidae: Dinopidae). Australian Zoologist 14: 189-197.
  • Coddington, J.A. (1986). Orb webs in 'non-orb' weaving ogre-faced spiders (Araneae: Dinopidae): a question of genealogy. Cladistics 2: 53.
  • Craig, L.C. and Bernard, G.D. (1990). Insect attraction to ultraviolet-reflecting spider webs and web decorations. Ecology 71 (2): 616-623.
  • Doran, N.E., Kiernan, K., Swain, R. and Richardson, A.M.M. (1999). Hickmania troglodytes, the Tasmanian cave spider, and its potential role in cave management. Journal of Insect Conservation 3: 257-262.
  • Doran, N.E., Richardson, A.M.M. and Swain, R. (1999). The Biology of Hickmania troglodytes, the Tasmanian Cave Spider. Pp. 330-332. In The Other 99%. The Conservation and Biodiversity of Invertebrates. W. Ponder and D. Lunney (eds). Transactions of the Royal Society of New South Wales.
  • Doran, N.E., Richardson, A.M.M. and Swain, R. The reproductive behaviour of the Tasmanian cave spider, Hickmania troglodytes (Araneae: Austrochilidae). Zoological Society of London 253: 405-418.
  • Downes, M.F. (1993). The life history of Badumna candida (Araneae, Amaurobioidea). Australian Journal of Zoology 41(5): 441-446.
  • Downes, M.F. (1994a). Tolerance, interattraction and cooperation in the behaviour of the social spider, Phryganoporus candidus (Araneae, Desidae). Bulletin of the British Arachnological Society 9(9): 309-317.
  • Downes, M.F. (1994b). Nest of the social spider Phryganoporus candidus (Araneae: Desidae): Structure, annual growth cycle and host plant relationships. Australian Journal of Zoology 42(2): 237-260.
  • Eisner, T. and Nowicki, S. (1983). Spider web protection through visual advertisement: role of the stabilimentum. Science 219: 185-187.
  • Elgar, M. and Fahey, B. (1966). Sexual canniballism, competition and size dimorphism in the orb-weaving spider Nephila plumipes Latreille (Araneae: Araneoidea). Behavioural Ecology 7: 195-198.
  • Elgar, M.A. (1991). Sexual cannibalism, size dimorphism and courtship behaviour in orb-weaving spiders (Araneidae). Evolution 45 (2): 444-448.
  • Elgar, M.A., Allen, R.A. and Evans, T.A. (1996). Foraging strategies in orb-spinning spiders: Ambient light and silk decorations in Argiope aetherea Walckenaer (Araneae: Araneoidea). Australian Journal of Ecology 21: 464-467.
  • Faulder, R.J. (1995). Systematics and biogeography of the spider genus Missulena Walckaener. Thesis (M.Sc.Agr.), University of Sydney.
  • Forster, L.M. (1995). The behavioural ecology of Latrodectus hasselti (Thorell) (Araneae: Theridiidae), the Australian Redback Spider: a review. In Australasian spiders and their relatives: papers honouring Barbara York Main (M.S. Harvey, ed.). Records of the Western Australian Museum Supplement 52: 13-24.
  • Forster, R.R., Platnick, N.I. and Gray, M.R. (1987). A review of the spider Superfamilies Hypochiloidea and Austrochiloidea (Araneae, Araneomorphae). Bulletin of the American Museum of Natural History 185 (1):1-116.
  • Gray, M. (1978). Silk, Spinnerets and Snares. Australian Natural History 19: 226-230.
  • Gray, M.R. (1983). The male of Progradungula carraiensis Forster and Gray (Araneae, Gradungulidae) with observations on the web and prey capture. Proceedings of the Linnean Society of New South Wales 107: 51-58.
  • Gray, M.R. (1987). Distribution of the funnelweb spiders. In Covacevitch, J., Davie, P. and Pearn, J. (eds), Toxic Plants and Animals. A Guide for Australia. Queensland Museum, Brisbane.
  • Gray, M.R. (1992). Funnelwebs: separating fact from fiction. Australian Natural History 24: 34-39.
  • Gray, M.R. (2002). The Taxonomy and Distribution of the Spider Genus Phryganoporus Simon (Araneae: Amaurobioidea: Desidae). Records of the Australian Museum (in press).
  • Gray, M.R. and Anderson, G.J. (1988). A new Australian species of Argyrodes (Araneoidea: Theridiidae) which preys upon its host. Proceedings of the Linnean Society of New South Wales 111(1): 25-30.
  • Herberstein, M.E. and Elgar, M.A. (1994). Foraging strategies of Eriophora transmarina and Nephila plumipes (Araneae: Araneoidea): Nocturnal and diurnal orb-weaving spiders. Australian Journal of Ecology 19: 451-457.
  • Hickman, V.V. (1967). Some Common Spiders of Tasmania. Tasmanian Museum and Art Gallery.
  • Isbister,G.K. & Gray, M.R. (2000). Acute and recurrent skin ulceration after spider bite. Medical Journal of Australia 172: 303-304.
  • Isbister, G.K. and Gray, M.R. (2002). A prospective study of 750 definite spider bites, with expert spider identification. Queensland Journal of Medicine ; 95: 723-731.
  • Isbister, G.K. and Gray, M.R. (2003). Latrodectism: a prospective cohort study of bites by formally identified redback spiders. Medical Journal of Australia , 179 (2): 88-91.
  • Isbister, G.K. and Gray, M.R. (2003). White-tail spider bite: a prospective study of 130 definite bites by Lampona species. Medical Journal of Australia , 179 (4): 199-202.
  • Jackson, R.R. (1996). Portia Spiders: Mistress of Deception. National Geographic 190 (5): 104-115.
  • Jackson, R.R. and Blest, A.D. (1982). The biology of Portia fimbriata, a web-building jumping spider (Araneae: Salticidae) from Queensland: utilisation of webs and predatory versatility. Journal of Zoology, London 196: 255-270.
  • Longman, H.A. (1922). The magnificent spider: Dicrostichus magnificus Rainbow: Notes on cocoon spinning and methods of catching prey. Proceedings of the Royal Society of Queensland 33: 91-98.
  • Main, B.Y. & Mascord, R. (1974). Description and natural history of a 'tube building' species of Dyarcyops from New South Wales and Queensland (Mygalomorphae: Ctenizidae). Journal of the Entomological Society of Australia (New South Wales) 8: 15-21.
  • Main, B.Y. (1953). Observations on the burrow and natural history of the trap-door spider Missulena (Ctenizidae). Western Australian Naturalist 5 (4): 73-80.
  • Main, B.Y. (1957). Biology of aganippine trapdoor spiders (Mygalomorphae: Ctenizidae). Australian Journal of Zoology 5 (4): 402-473.
  • Main, B.Y. (1973). Spiders. Australian Naturalists Library. Collins, Sydney.
  • Main, B.Y. (1976). Spiders. The Australian Naturalist Library: Collins, Sydney.
  • Main, B.Y. (1982). Adaptations to arid habitats by mygalomorph spiders. pp 273-283. In W.R. Barker and P.J.M. Greenslade (eds). Evolution of the Flora and Fauna of Arid Australia. Peacock Publishers, South Australia.
  • McGhee, K. (1999). Funnel-webs. Australian Geographic 53: 80-95.
  • McKeown, K. (1963). Australian Spiders. Angus and Robertson, Sydney.
  • Platnick, N.I. (2000). A relimitation and revision of the Australasian ground spider family Lamponidae (Araneae: Gnaphosoidea). Bulletin of the American Museum of Natural History 245: 330 pp.
  • Raven R. and J. Gallon (1987). The Redback Spider. In Covacevich, J., Davie, P., Pearn, J. (eds), Toxic Plants and Animals: a guide for Australia. Queensland Museum.
  • Robins, M. and Le Page, M. (2000). Mean and green - Does a deadly spider hold the key to eco-pesticides? New Scientist, 17 June, No. 2243: 5.
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